Separation for Aidan resulted in Grayson showing dominance over Denali.  Aidan and the arctics were separated from Denali and Boltz by two hours while Aidan recovered from his surgery. Grayson met Denali at the chute and dominated him immediately upon Denali’s entry back into the exhibit. Separation within a wolf pack can create a re-assertion of rank order, and as a young wolf, Grayson is tuned in to opportunities to increase his rank. Grayson continues to periodically give a solo, haunting howl. He seems to do this when he is keyed in to Aidan’s moods and/or stress.