The secrets in scat
From The Lewiston Tribune in Montana:
A recent study determined that hunting and trapping can be an effective wolf management tool, and animals from nearby packs don’t necessarily join depleted packs to compensate for the harvest.
The work published in the journal Animal Conservation looked at wolf populations in central Idaho before and after hunting started in 2011, and at populations in southern Alberta, Canada, that have been heavily hunted for decades. Researchers, including scientists from the University of Idaho and the University of Montana, assumed packs reduced by harvest would quickly rebound by accepting immigrants and that it would happen more frequently in Alberta because of the higher harvest levels there.