From sci-news.com:

The scientists used a range of genetic techniques to extract and examine DNA taken from wolf and dog fur samples as well as wolf scat and blood samples. They found recent hybrid ancestry in about ten percent of the dogs and wolves sampled.

About 2-3 percent of the sampled wolves and dogs were identified as first-generation hybrids. This included hybridization between wolves and the shepherd dogs used to guard sheep from wolf attacks.

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